Dactylorhiza

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Dactylorhiza
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Common Spotted Orchid (Dactylorhiza fuchsii)
Common Spotted Orchid
(Dactylorhiza fuchsii)
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Order: Asparagales
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Family: Orchidaceae
Subfamily: Orchidoideae
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Tribe: Orchideae
Subtribe: Orchidinae
Genus: Dactylorhiza
Necker ex Nevski
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Species
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Dactylorhiza Necker ex Nevski 1937, is a genus of terrestrial (ground-dwelling) plants from the orchid family (Orchidaceae).

The name Dactylorhiza is derived from Greek words δάκτυλος "daktylos" (finger) and ρίζα "rhiza" (root). This is because of the shape of the genus' two underground tubers. Dactylorhiza were previously classified under Orchis.

These orchids are distributed throughout the subarctic and temperate northern hemisphere : in Europe, from Scandinavia to North Africa; also on Madeira, Iceland, West Asia, North Asia, the Himalayas, North America and even in Alaska.

These terrestrial orchids grow in basic soils in wet meadows, bogs, heathland and in areas sparsely populated by trees. They are tuberous geophytes. In a thickened underground stem, they can store a large amount of water to survive arid conditions.

The long leaves are lanceolate and, in most species, also speckled. They grow along a rather long stem which reaches a height of 70-90 cm. Leaves higher on the stem are shorter than leaves lower on the stem.

The inflorescence, compared to the length of the plant, is rather short. It consists of a compact raceme with 25-50 flowers. These develop from axillary buds. The dominant colors are all shades of pink to red, sprinkled with darker speckles.

Many 'species' in this genus hybridise so readily that species boundaries themselves are vague, with regular name changes and no clear answers. A few species colonise very well onto fresh industrial wastes such as pulverised fuel ash, where vast hybrid swarms can appear for a decade or more, before ecological succession replaces them.

Contents

Species

Spotted Orchid
(Dactylorhiza incarnata)
Leopard Marsh Orchid
(Dactylorhiza praetermissa)
Broad-leaved Marsh Orchid
(Dactylorhiza majalis)

There are many species that cross-fertilize, giving an immense variation and complicating classification.

  • Dactylorhiza alpestris : Alpine Dactylorhiza (Pyrenees, Alps, Carpathians).
  • Dactylorhiza angustata (France).
  • Dactylorhiza aristata : Keyflower (E. China to Alaska).
    • Dactylorhiza aristata var. aristata : Keyflower (E. China to Alaska). Tuber geophyte
    • Dactylorhiza aristata var. kodiakensis : Kodiak Keyflower (Aleutian Is. to SW. Alaska). Tuber geophyte
  • Dactylorhiza armeniaca (Turkey) - has become synonym of Dactylorhiza euxina subsp. armeniaca (Hedrén) Kreutz
  • Dactylorhiza atlantica Kreutz & Vlaciha (Morocco)
  • Dactylorhiza baldshuanica (C. Asia).
  • Dactylorhiza baltica (Eastern Europe) (synonym of Dactylorhiza longifolia (Neuman) Aver.)
  • Dactylorhiza baumanniana (N. Greece).
    • Dactylorhiza baumanniana subsp. smolikana (B. Willing & E. Willing) H. Baumann & R. Lorenz (Greece)
  • Dactylorhiza bohemica (EC. Europe).
  • Dactylorhiza cordigera (Fr.) Soó (SE. Europe to Ukraine).
    • Dactylorhiza cordigera subsp. bosniaca (N. Balkan Pen). Tuber geophyte
    • Dactylorhiza cordigera subsp. cordigera (SE. Europe to Ukraine).
    • Dactylorhiza cordigera var. graeca (H.Baumann) Presser)
    • Dactylorhiza cordigera subsp. pindica (B. Willing & E. Willing) H. Baumann & R. Lorenz (NW. Greece).
    • Dactylorhiza cordigera var. rhodopeia Presser (Greece, Southeastern Europe, Europe)
    • Dactylorhiza cordigera subsp. siculorum (Romania to W. Ukraine). Tuber geophyte
  • Dactylorhiza elata (Poir.) Soó : Stately Dactylorhiza (W. Europe to NW. Africa).
    • Dactylorhiza elata subsp. ambigua (Martrin-Donos) Kreutz
    • Dactylorhiza elata subsp. brennensis (W. Europe). Tuber geophyte
    • Dactylorhiza elata subsp. elata (NW. Africa. Tuber geophyte
    • Dactylorhiza elata subsp. mauritanica B.Baumann & H. Baumann (Morocco, Algeria)
    • Dactylorhiza elata subsp. sesquipedalis (SW. Europe to Sicilia). Tuber geophyte
  • Dactylorhiza euxina (Nevski) Czerep.
    • Dactylorhiza euxina subsp. armeniaca (Hedrén) Kreutz
  • Dactylorhiza flavescens (Turkey to C. Asia).
  • Dactylorhiza foliosa : Richly-leaved Dactylorhiza (Madeira).
  • Dactylorhiza fuchsii (Druce) Soó  : Common Spotted Orchid, Fuch's Dactylorhiza (Europe to Siberia).
    • Dactylorhiza fuchsii subsp. carpatica (Batousek & Kreutz) Kreutz
    • Dactylorhiza fuchsii subsp. fuchsii (Europe to Siberia).
    • Dactylorhiza fuchsii subsp. hebridensis (W. Europe).
    • Dactylorhiza fuchsii subsp. meyeri (Rchb.f.) Kulikov & E.G.Philippov
    • Dactylorhiza fuchsii subsp. okellyi (Ireland, W. Great Britain).
    • Dactylorhiza fuchsii subsp. psychrophila (Europe to Siberia).
    • Dactylorhiza fuchsii var. sooana (Borsos) Kreutz (Hungary)
    • Dactylorhiza fuchsii var. sudetica (Poech ex Rchb.f.) H.Baumann
  • Dactylorhiza gervasiana (Sicilia to S. Italy).
  • Dactylorhiza graeca (N. Greece) - has become synonym of Dactylorhiza cordigera var. graeca (H.Baumann) Presser)
  • Dactylorhiza graggeriana (W. Himalaya).
  • Dactylorhiza hatagirea (Pakistan to SE. Tibet).
  • Dactylorhiza iberica (Greece to Iran).
  • Dactylorhiza ilgazica (N. Turkey) - now synonym of Dactylorhiza urvilleana subsp. ilgazica (Kreutz) Kreutz
  • Dactylorhiza incarnata (L.) Soó  : Spotted Orchid (Europe to Mongolia).
    • Dactylorhiza incarnata var. baumgartneriana (B.Baumann, H.Baumann, R.Lorenz & Ruedi Peter) P.Delforge
    • Dactylorhiza incarnata subsp. coccinea : Early Marsh Orchid, Northern Marsh Orchid (Great Britain, Ireland). Tuber geophyte
    • Dactylorhiza incarnata subsp. cruenta (Europe to Turkey). Tuber geophyte
    • Dactylorhiza incarnata subsp. gemmana (W. Europe). Tuber geophyte
    • Dactylorhiza incarnata subsp. incarnata (Europe to Mongolia). Tuber geophyte
    • Dactylorhiza incarnata nothosubsp. krylovii (W. Europe to Siberia).Tuber geophyte
    • Dactylorhiza incarnata subsp. lobelii (Norway to The Netherlands). Tuber geophyte
    • Dactylorhiza incarnata subsp. ochroleuca (Europe). Tuber geophyte
    • Dactylorhiza incarnata subsp. pulchella (Europe). Tuber geophyte
    • Dactylorhiza incarnata nothosubsp. versicolor (Europe) Tuber geophyte
  • Dactylorhiza insularis : Island Dactylorhiza (W. Medit. to WC. Italy).
  • Dactylorhiza kafiriana (NE. Afghanistan to W. Himalaya).
  • Dactylorhiza kalopissii E.Nelson (N. Greece).
    • Dactylorhiza kalopissii subsp. macedonica (J.Hölzinger & Künkele) Kreutz
    • Dactylorhiza kalopissii subsp. pythagorae (Gölz & H.R.Reinhard) Kreutz
  • Dactylorhiza kulikalonica (C. Asia).
  • Dactylorhiza lapponica (Laest.ex Hartm.) Soó (N. Europe).
    • Dactylorhiza lapponica subsp. angustata (Arv.-Touv.) Kreutz
    • Dactylorhiza lapponica subsp. rhaetica H. Baumann & R. Lorenz (Alps of Austria, Germany, Switzerland, Italy and France)
  • Dactylorhiza lapponica subsp. russowii (Klinge) H. Baumann & R. Lorenz (C. Europe to Siberia).
  • Dactylorhiza libanotica (Lebanon)
  • Dactylorhiza longifolia (Europe to C. Asia).
  • Dactylorhiza macedonica (N. Greece) - now a synonym of Dactylorhiza kalopissii subsp. macedonica (J.Hölzinger & Künkele) Kreutz
  • Dactylorhiza maculata (L.) Soó : Heath Spotted Orchid, Moorland Spotted Orchid (NW. Africa, Europe to Siberia).
    • Dactylorhiza maculata subsp. battandieri (N. Algeria).
    • Dactylorhiza maculata subsp. caramulensis (W. Europe).
    • Dactylorhiza maculata subsp. elodes (Europe).
    • Dactylorhiza maculata subsp. ericetorum : Heath Spotted Orchid (W. Europe).
    • Dactylorhiza maculata subsp. islandica (Iceland).
    • Dactylorhiza maculata subsp. kolaensis (Montell) Kreutz
    • Dactylorhiza maculata subsp. maculata (Europe to Siberia).
    • Dactylorhiza maculata subsp. maurusia (Morocco)
    • Dactylorhiza maculata subsp. podesta (Netherlands).
    • Dactylorhiza maculata subsp. rhoumensis (Great Britain)
    • Dactylorhiza maculata subsp. savogiensis (D.Tyteca & Gathoye) Kreutz
    • Dactylorhiza maculata subsp. schurii (Carpathians).
    • Dactylorhiza maculata subsp. sennia (Vollmar) Kreutz
    • Dactylorhiza maculata subsp. transsilvanica (SC. & SE. Europe).
  • Dactylorhiza magna (C. Asia).
  • Dactylorhiza majalis (Rchb.) P.F.Hunt & Summerh.  : Broad-leaved Marsh Orchid, Western Marsh Orchid, Fan Orchid, Common Marsh Orchid (Europe).
    • Dactylorhiza majalis var. brevifolia (Rchb.f.) Kreutz
    • Dactylorhiza majalis subsp. calcifugiens (Denmark)
    • Dactylorhiza majalis subsp. cambrensis (Coastal Great Britain and Denmark) (now synonym of Dactylorhiza purpurella (T.Stephenson & T.A.Stephenson) Soó var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm 2005)
    • Dactylorhiza majalis subsp. majalis (Europe). Tuber geophyte
    • Dactylorhiza majalis subsp. occidentalis (W. & SW. Ireland, N. Great Britain).
    • Dactylorhiza majalis subsp. parvimajalis (D.Tyteca & Gathoye) Kreutz
    • Dactylorhiza majalis subsp. purpurella : Early Marsh Orchid, Northern Marsh Orchid (NW. Europe).
    • Dactylorhiza majalis subsp. sphagnicola
    • Dactylorhiza majalis subsp. turfosa (Alps to W. Carpathians) - has become a synonym of Dactylorhiza traunsteineri subsp. turfosa (F.Proch.) Kreutz
  • Dactylorhiza markusii : Markus' Dactylorhiza (N. Portugal to W. Spain and Italy).
  • Dactylorhiza nieschalkiorum (N. Turkey).
  • Dactylorhiza osmanica (Turkey to Syria).
    • Dactylorhiza osmanica var. anatolica (Turkey). Tuber geophyte
    • Dactylorhiza osmanica var. osmanica (Turkey to Syria). Tuber geophyte
  • Dactylorhiza pindica (NW. Greece).
  • Dactylorhiza praetermissa : Leopard Marsh Orchid, Southern Marsh Orchid (W. & NW. Europe)
    • Dactylorhiza purpurella (T.Stephenson & T.A.Stephenson) Soó var. cambrensis (R.H.Roberts) R.M.Bateman & Denholm 2005
  • Dactylorhiza pythagorae (E. Aegean Is.) - now a synonym of Dactylorhiza kalopissii subsp. pythagorae (Gölz & H.R.Reinhard) Kreutz
  • Dactylorhiza romana : Roman Dactylorhiza (Mediterranean)
  • Dactylorhiza saccifera (Brongn.) Soó  : Sack-carrying Dactylorhiza (Mediterranean).
    • Dactylorhiza saccifera subsp. bithynica (H.Baumann) Kreutz
    • Dactylorhiza saccifera subsp. gervasiana (Tod.) Kreutz
  • Dactylorhiza salina (Caucasus to Amur)
  • Dactylorhiza sambucina : Elder-flowered Orchid (Europe). Photos
  • Dactylorhiza sudetica (Europe to Siberia)
  • Dactylorhiza traunsteineri (Saut. ex Rchb.) Soó : Narrow-leaved Marsh Orchid, Traunstein's Dactylorhiza, Irish Marsh Dactylorhiza (Europe to W. Siberia).
    • Dactylorhiza traunsteineri subsp. carpatica (Slovakia) - has become synonym of Dactylorhiza fuchsii subsp. carpatica (Batousek & Kreutz) Kreutz
    • Dactylorhiza traunsteineri subsp. turfosa (F.Proch.) Kreutz
    • Dactylorhiza traunsteineri subsp. 'curvifolia' (N. & NE. Europe).
    • Dactylorhiza traunsteineri subsp. traunsteineri (Europe to W. Siberia).
    • Dactylorhiza traunsteineri subsp. wirtgenii (Höppner) Kreutz
    • Dactylorhiza traunsteinerioides (Pugsley)Landwehr (synonym of Dactylorhiza traunsteineri subsp. traunsteineri)
  • Dactylorhiza umbrosa (W. & C. Asia to Siberia)
  • Dactylorhiza urvilleana (Steud.) H.Baumann & Künkele (N. & NE. Turkey to Iran)
    • Dactylorhiza urvilleana subsp. bithynica (H.Baumann) H. Baumann & R. Lorenz
    • Dactylorhiza urvilleana subsp. phoenissa B. Baumann & H. Baumann (Lebanon)
    • Dactylorhiza urvilleana subsp. ilgazica (Kreutz) Kreutz
  • Dactylorhiza viridis (Subarctic and subalpine Northern Hemisphere).
    • Dactylorhiza viridis var. virescens (Temp. Asia, N. America)
    • Dactylorhiza viridis var. viridis (Subarctic and subalpine Northern Hemisphere)

Hybrids

Dactylorhiza × aschersoniana
Dactylorhiza × braunii

Note : nothosubspecies = a hybrid subspecies; nothovarietas = subvariety.

  • Dactylorhiza × abantiana (D. iberica × D. nieschalkiorum) (Turkey).
  • Dactylorhiza × aldenii (D. iberica × D. kalopissii) (Greece).
  • Dactylorhiza × altobracensis (D. maculata × D. sambucina) (France, Austria).
  • Dactylorhiza × aschersoniana (D. incarnata × D. majalis) (W. & C. Europe).
    • Dactylorhiza × aschersoniana nothosubsp. aschersoniana (W. & C. Europe). Tuber geophyte
    • Dactylorhiza × aschersoniana nothosubsp. templinensis (D. incarnata subsp. ochroleuca × D. majalis) (C. Europe). Tuber geophyte
    • Dactylorhiza × aschersoniana nothovar. uliginosa (D. incarnata subsp.pulchella × D. majalis) (C. Europe). Tuber geophyte
  • Dactylorhiza × baicalica (D. incarnata subsp. cruenta × D. salina) (Siberia).
  • Dactylorhiza × balabaniana (D. iberica × D. urvilleana) (Turkey).
  • Dactylorhiza × bayburtiana (D. euxina × D. umbrosa) (Turkey).
  • Dactylorhiza × beckeriana (C. Europe).
  • Dactylorhiza × boluiana (D. nieschalkiorum × D. saccifera) (Turkey).
  • Dactylorhiza × bourdonii (D. brennensis × D. incarnata) (France).
  • Dactylorhiza × braunii (D. fuchsii × D. majalis) (Europe).
    • Dactylorhiza × braunii nothosubsp. braunii (Europe). Tuber geophyte
    • Dactylorhiza × braunii nothosubsp. lilacina (D fuchsii × D. majalis subsp. turfosa) (EC. Europe). Tuber geophyte
    • Dactylorhiza × braunii nothosubsp. monticola (D. fuchsii subsp. psychrophila × D. majalis) (Europe). Tuber geophyte
    • Dactylorhiza × braunii nothosubsp. smitakii (D. fuchsii subsp. sooana × D. majalis) (EC. Europe). Tuber geophyte
  • Dactylorhiza × breviceras (D. osmanica × D. urvilleana) (Turkey).
  • Dactylorhiza × carnea (D. incarnata × D. maculata subsp. ericetorum) (W. Europe).
    • Dactylorhiza × carnea nothosubsp. ampolai (D. incarnata subsp. cruenta × D. maculata) (Europe). Tuber geophyte
    • Dactylorhiza × carnea nothosubsp. carnea (W. Europe). Tuber geophyte
    • Dactylorhiza × carnea nothosubsp. maculatiformis. (D. incarnata × D. maculata) (W. Europe). Tuber geophyte
  • Dactylorhiza × claudiopolitana (D. incarnata × D. schurii) (Europe.
  • Dactylorhiza × conigerum (D. maculata × D. viridis) (W. Europe).
  • Dactylorhiza × csatoi (D. cordigera × D. maculata) (SE. Europe).
  • Dactylorhiza czerniakowskae (C. Asia).
  • Dactylorhiza × daunia (D. romana × D. saccifera) (S. Europe).
  • Dactylorhiza × delamainii (D. elata subsp. sesquipedalis × D. maculata) (SW. Europe).
  • Dactylorhiza × dinglensis (D. maculata subsp. ericetorum × D. majalis subsp. occidentalis) (W. Europe).
    • Dactylorhiza × dinglensis nothosubsp. dinglensis (W. Europe). Tuber geophyte
    • Dactylorhiza × dinglensis nothosubsp. robertsii (D. maculata subsp. ericetorum × D. majalis subsp. cambrensi) (Great Britain). Tuber geophyte
    • Dactylorhiza × dinglensis nothosubsp. senayi (D. maculata subsp. elodes × D. majalis) (Europe).Tuber geophyte
    • Dactylorhiza × dinglensis nothosubsp. townsendiana (D. maculata subsp. ericetorum × D. majalis) (Europe). Tuber geophyte
    • Dactylorhiza × dinglensis nothosubsp. vermeuleniana (D. maculata × D. majalis) (W. Europe). Tuber geophyte
  • Dactylorhiza × drucei (D. majalis × D. viridis) (W. Europe)
  • Dactylorhiza × dubreuilhii (D. elata subsp. sesquipedalis × D. incarnata) (W. Europe).
  • Dactylorhiza × dufftiana (D. majalis × D. traunsteineri) (Europe).
  • Dactylorhiza × dufftii (D. incarnata × D. traunsteineri) (Europe).
    • Dactylorhiza × dufftii nothosubsp. dufftii (Europe). Tuber geophyte
    • Dactylorhiza × dufftii nothosubsp. gotlandica (D incarnata subsp. ochroleuca × D. traunsteineri) (Europe). Tuber geophyte
    • Dactylorhiza × dufftii nothosubsp. stenkyrkae (D. incarnata subsp. cruenta × D. traunsteineri) (Europe). Tuber geophyte
  • Dactylorhiza × erdingeri (D. sambucina × D. viridis) (W. Europe).
  • Dactylorhiza euxina (NE. Turkey to Caucasus).
    • Dactylorhiza euxinavar. euxina (NE. Turkey to Caucasus). Tuber geophyte
    • Dactylorhiza euxinavar. markowitschii (NE. Turkey to Caucasus). Tuber geophyte
  • Dactylorhiza × flixensis (D. incarnata subsp. pulchella × D. traunsteineri.) (Switzerland).
  • Dactylorhiza × formosa (D. maculata subsp. ericetorum × D. purpurella) (W. Europe).
  • Dactylorhiza × fourkensis (D. baumanniana × D. sambucina) (Greece).
  • Dactylorhiza × gabretana (D. incarnata × D. maculata × D. sambucina) (Europe).
  • Dactylorhiza × genevensis (D. incarnata × D. latifolia × D. maculata) (Europe).
  • Dactylorhiza × godferyana (D. majalis × D. praetermissa) (W. Europe).
  • Dactylorhiza × grandis (D. fuchsii × D. praetermissa) (W. Europe).
  • Dactylorhiza × guilhotii (D. incarnata × D. viridis) (W. Europe).
  • Dactylorhiza × guillaumeae (D. incarnata × D. sambucina) (W. Europe).
  • Dactylorhiza × gustavssonii (D. iberica × D. saccifera) (Greece to Turkey).
  • Dactylorhiza × hallii (D maculata subsp. ericetorum × D. praetermissa) (W. Europe).
    • Dactylorhiza × hallii nothosubsp. hallii (W. Europe). Tuber geophyte
    • Dactylorhiza × hallii nothosubsp. nummiana (D. maculata subsp. elodes × D. praetermissa) (W. Europe). Tuber geophyte
  • Dactylorhiza × hochreutinerana (D. alpestris × D. incarnata) (W. Europe).
  • Dactylorhiza × insignis (D. praetermissa × D. purpurella) (W. Europe).
  • Dactylorhiza × ishorica (D. incarnata × D. longifolia) (European Russia).
  • Dactylorhiza × jenensis (D. maculata subsp. ericetorum × D. traunsteineri) (W. & NC. Europe)
  • Dactylorhiza × jestrebiensis (D. bohemica × D. majalis) (EC. Europe).
  • Dactylorhiza × juennensis (D. fuchsii × D. lapponica) (C. Europe).
  • Dactylorhiza × katarana (D kalopissii × D. saccifera) (Greece).
  • Dactylorhiza × kelleriana (D. fuchsii × D. traunsteineri) (Europe).
  • Dactylorhiza × kerasovinensis (D. pindica × D. saccifera) (Greece).
  • Dactylorhiza × kerneriorum (D. fuchsii × D. incarnata) (Europe).
    • Dactylorhiza × kerneriorum nothosubsp. kerneriorum (Europe). Tuber geophyte
    • Dactylorhiza × kerneriorum nothosubsp. lillsundica (D. fuchsii × D. incarnata subsp. ochroleuca) (N. & W. Europe). Tuber geophyte
    • Dactylorhiza × kerneriorum nothosubsp. variablis (D. fuchsii subsp. hebridensis × D. incarnata) (W. Europe). Tuber geophyte
  • Dactylorhiza × komiensis (D. hebridensis × D. maculata) (

E. Europe).

  • Dactylorhiza × kopdagiana (D. iberica × D. umbrosa) (Turkey).
  • Dactylorhiza × koutsourana (D. baumanniana × D. smolikana) (Greece).
  • Dactylorhiza × kuuskiae (D.longifolia × D. traunsteineri) (E. Europe).
  • Dactylorhiza × latirella (D. incarnata × D. purpurella) (W. Europe).
  • Dactylorhiza × lehmannii (D. incarnata × D. russowii) (Europe).
  • Dactylorhiza × megapolitana (D. fuchsii × D. russowii) (C. Europe).
  • Dactylorhiza × metsowonensis (D. kalopissii × D. sambucina) (Greece).
  • Dactylorhiza × mixtum (D. fuchsii × D. viridis) (W. Europe).
  • Dactylorhiza × mulignensis (D. incarnata subsp. pulchella × D. majalis) (C. Europe).
  • Dactylorhiza × nevskii (D. osmanica × D. umbrosa) (Turkey).
  • Dactylorhiza × ornonensis (D. elata subsp. sesquipedalis × D. incarnata × D. maculata) (W. Europe).
  • Dactylorhiza × paridaeniana (D. elata subsp. sesquipedalis × D. praetermissa) (W. Europe).
  • Dactylorhiza × pontica (D. urvilleana × D. viridis) (Turkey)
  • Dactylorhiza × prochazkana (D. bohemica × D. maculata) (EC. Europe).
  • Dactylorhiza × renzii (D. incarnata × D. nieschalkiorum) (Turkey).
  • Dactylorhiza × rizeana (D. euxina × D. urvilleana) (Turkey).
  • Dactylorhiza × rombucina (D. romana × D. sambucina) (C. Europe).
  • Dactylorhiza × ruppertii (D. majalis × D. sambucina) (Europe).
  • Dactylorhiza × salictina (D. pindica × D. smolikana) (Greece).
  • Dactylorhiza × serbica (D. incarnata × D. saccifera) (Europe).
  • Dactylorhiza × serreana (D. graeca × D. lagotis) (Greece).
  • Dactylorhiza × sivasiana (D. umbrosa × D. urvilleana) (Turkey).
  • Dactylorhiza × sooi (D. alpestris × D. fuchsii.) (Europe).
  • Dactylorhiza × souflikensis (D. baumanniana × D. pindica) (Greece).
  • Dactylorhiza × stagni-novi (D. brennensis × D. fuchsii) (Europe).
  • Dactylorhiza × szaboiana (D. cordigera × D. sudetica) (SE. Europe).
  • Dactylorhiza × transiens (D. fuchsii × D. maculata subsp. ericetorum) (Europe)
    • Dactylorhiza × transiens nothosubsp. corylensis (D. fuchsii subsp. hebridensis × D. maculata)
    • Dactylorhiza × transienssubsp. ericetorum (Europe). Tuber geophyte
    • Dactylorhiza × transiens nothosubsp. transiens (Europe). Tuber geophyte
  • Dactylorhiza × turcestanicum (D. umbrosa × D. viridis) (C. Asia).
  • Dactylorhiza × vallis-peenae (D. majalis × D. russowii) (C. Europe).
  • Dactylorhiza × venusta (D. fuchsii × D. purpurella) (Europe).
    • Dactylorhiza × venusta nothosubsp. hebridella (D. fuchsii subsp. hebridensis × D. purpurella) (Great Britain). Tuber geophyte
    • Dactylorhiza × venusta nothosubsp. venusta (Europe) Tuber geophyte
  • Dactylorhiza × viridella (D. purpurella × D. viridis) (W. Europe).
  • Dactylorhiza × vitosana (D. saccifera × D. sambucina) (SE. Europe).
  • Dactylorhiza × vogtiana (D. iberica × D. incarnata) (Turkey).
  • Dactylorhiza × vorasica (D. cordigera × D. sambucina) (Greece).
  • Dactylorhiza × weissenbachiana (D. incarnata × D. lapponica) (C. Europe).
  • Dactylorhiza × wiefelspuetziana (D. maculata × D. sphagnicola) (W. Europe).

This list follows the World checklist of monocotyledons,[1] periodically amended from the "Orchid Research Newsletter", both of Kew Botanical Gardens.[2]

Notes

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External links

  • [htt

p://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=109214 Flora of North America Genus page]

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